Morphological variation and asymmetrical development in the skull of Styracosaurus albertensis

The ceratopsid Styracosaurus albertensis is an important component of the herbivorous dinosaur fauna preserved in the Campanian-aged Dinosaur Park Formation in Alberta, Canada. A new skull (UALVP55900) exhibits previously unobserved variability and asymmetry in this species. As in the type, the right lateral parietal bar bears seven epiossifications, but the left parietal bar has eight. Epiossifications p3‒p6 are asymmetrical with respect to...