Butterflies in Swedish grasslands benefit from forest and respond to landscape composition at different spatial scales

Loss and fragmentation of semi-natural grasslands has critically affected many butterfly species in Europe. Habitat area and isolation can have strong effects on the local biodiversity but species may also be strongly affected by the surrounding matrix. We explored how different land cover types in the landscape explained the occurrence of butterfly species in semi-natural grasslands. Using data from 476 semi-natural grasslands in Sweden, we...