Limuloid trackways from Permian-Triassic continental successions of North China

Forty-four well-preserved specimens of Kouphichnium, a typical limuloid (horseshoe crab) trackway, are reported and systematically studied for the first time from continental Upper Permian to Lower Triassic successions in North China. A new ichnotaxonomic system of Kouphichnium is proposed according to the ichnotaxobase of limuloid trackways, including simple foot imprints (T), pusher imprints (P), median impressions (M) and genal spine...