Reinterpretation of a previously described Jehol bird clarifies early trophic evolution in the Ornithuromorpha

STM35-3 from the Yixian Formation is the only Early Cretaceous ornithuromorph preserving direct evidence of granivory. The crop contains numerous seeds and the preservation of gastroliths presumably within the ventriculus indicates this diet was paired with the presence of a gastric mill as in living granivorous birds. STM35-3 was originally referred to Hongshanornis longicresta , member of a diverse clade of small, basal ornithuromorphs with...